Sue and Will become unlikely allies when Principal Figgins hires Shannon Beiste to coach the football team and cuts the Cheerios and New Directions' budgets. But when Sue's plot against Coach Beiste backfires, she becomes even more determined to take down the glee club.

Sue continues to show a softer side in her relationships with her sister and her Cheerios sidekick, Becky, who both have Down syndrome. Still, she doesn't stop waging attacks on Will. And when Sue becomes acting principal, she uses her new authority to try to disband the football team.

After a go at online dating, Sue realizes that her only perfect match is, in fact, Sue Sylvester. So she decides to get married to herself.

Sue comes up with a plan for the most exciting Cheerios routine ever, which involves firing Brittany from a cannon. When Brittany refuses, Quinn and Santana quit the Cheerios with her in protest. The squad is defeated at nationals in what would have been Sue's seventh consecutive title, and the team's budget shifts to the glee club.

Now more vindictive than ever, Sue amps up her plots against New Directions by becoming coach of rival show choir Aural Intensity; she also tells the competing Warblers that New Directions is "turning up the sexy" at regionals. Then after the McKinley glee club's win at the competition, she decides to create a "League of Evil" to destroy them once and for all.

When Sue's beloved sister, Jean, unexpectedly dies, the New Directions team creates a "Willy Wonka"-themed funeral on her behalf. Sue is touched and vows to stop trying to destroy the glee club, as she moves on to a new mission: running for a seat in the U.S. House of Representatives.

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Jane Lynch cut her theatrical teeth at The Second City, Steppenwolf Theatre and in many church basements all over the greater Chicago area. This year, Lynch won an Emmy Award for Outstanding Supporting Actress in a Comedy Series for her work on GLEE.

Lynch's film credits include "Julie & Julia," "Shrek Forever After," "Post Grad" and "Paul and Brownie Masters." Her past film work includes Christopher Guest's "For Your Consideration," "A Mighty Wind" and "Best in Show," as well as "Role Models," "The Rocker," "Spring Breakdown," "Space Chimps," "Alvin and the Chipmunks," "Walk Hard: The Dewey Cox Story," "Talladega Nights: The Ballad of Ricky Bobby," "The 40 Year Old Virgin," Margaret Cho's "Celeste and Bam Bam," Alan Cumming's "Suffering Man's Charity," "Lemony Snicket's A Series of Unfortunate Events," "Sleepover" and "Surviving Eden." She recently wrapped production on "The Three Stooges" film.

Her recent television credits include "Party Down," "Lovespring International," "Desperate Housewives," "Weeds" and "The L Word." Lynch has had recurring roles on "Boston Legal," "Criminal Minds," "The New Adventures of Old Christine" and "Two and a Half Men," for which she received an Emmy nomination for Outstanding Guest Actress in a Comedy Series.

Lynch's play "Oh Sister, My Sister!" had runs at the Tamarind Theatre and Bang Theater, garnering the LA Weekly Comedy Ensemble of the Year Award.

She lives in Los Angeles.